Budgie Genetics Guide
Research date: April 8, 2026
This document has been prepared to collect budgie color genetics and mutations in one place. The goal is to answer the question "What happens when genetics match with which genetics?" with a rule-based system rather than a memorized list.
Scope
- This guide lists all the mutations supported in the project.
- It does not enumerate all possible multiple combinations; instead, it gives the inheritance type, locus logic, and frequent combinations.
- If more than one independent locus is located in the same offspring, the total probability is calculated by multiplying the single locus probabilities.
- Since crossing-over can be seen in close genes on the Z chromosome, there may be small differences between theoretical rates and practical results. This situation is the most
opaline,cinnamon,inoAndslateIt is important for. [K6] [K12]
Quick Dictionary
visual: The bird visibly carries the mutation.splitorcarrier: The bird carries the mutation without displaying it in the phenotype.AR: Autosomal recessive.AD: Autosomal dominant.AID: Autosomal incomplete dominant / single factor-double factor mutation.SLR: Sex-linked recessive.SF: Single factor, single factor.DF: Double factor, double factor.
Genetic Map
Basic tones in WBO color standards Light Green, Dark Green, Olive Green, Grey Green, Skyblue, Cobalt, Mauve, Violet And Grey is given as . [K1] [K2]
Basic Inheritance Rules
1. Autosomal recessive mutations
This is the rule blue, dilute, greywing, clearwing, recessive_pied, clearflight_pied, fallow types, saddleback, faded, mottled and similar mutations.
| matching | Conclusion |
|---|---|
| visual x visual | %100 visual |
| visual x split | %50 visual, %50 split |
| visual x normal | %100 split |
| split x split | %25 visual, %50 split, %25 normal |
| split x normal | %50 split, %50 normal |
| normal x normal | %100 normal |
2. Autosomal dominant mutations
This is the rule grey, dominant_pied, dominant_clearbody, crested and you use it in some controversial modern mutations.
| matching | Conclusion |
|---|---|
| mutant x normal | Approximately %50 mutant, %50 normal |
| mutant x mutant | Approximately %75 mutant, %25 normal |
Note: In dominant mutations, heterozygous and homozygous individuals sometimes do not look the same. crested and health/expression differences are significant in some area types. [K10]
3. Autosomal incomplete dominant mutations
This is the rule dark_factor, violet, spangle, anthracite and you use it on some face alleles at the blue-locus.
| matching | Conclusion |
|---|---|
| SF x normal | %50 SF, %50 normal |
| SF x SF | %25 DF, %50 SF, %25 normal |
| DF x normal | %100 SF |
| DF x SF | %50 DF, %50 SF |
| DF x DF | %100 DF |
4. Sex-linked recessive mutations
male budgie ZZ, female ZW is accepted. female birds split I can't be; If they inherit the mutant allele, they become visible.
This is the rule ino, pallid, opaline, cinnamon, slate, texas_clearbody You can use it in Z-linked mutations and so on.
| matching | male puppies | female offspring |
|---|---|---|
erkek visual x disi normal |
%100 split |
%100 visual |
erkek split x disi normal |
%50 split, %50 normal |
%50 visual, %50 normal |
erkek normal x disi visual |
%100 split |
%100 normal |
erkek visual x disi visual |
%100 visual |
%100 visual |
erkek split x disi visual |
%50 visual, %50 split |
%50 visual, %50 normal |
Z Chromosome and Crossing-Over
Approximate crossing-over values according to MUTAVI sources:
opaline <-> cinnamon: approximately%32[P6]opaline <-> slate: approximately%40.5[P6]cinnamon <-> slate: approximately%5 veya daha az[K6] [K11]cinnamon <-> ino: approximately%3[K12]
This is especially lacewing, opaline-cinnamon, cinnamon-slate And opaline-slate It means that their matings may result in unexpected but biologically explainable offspring. [K6] [K12]
Special Series Working at the Same Locus
1. Blue locus
Many variations affecting the blue series and yellow/face are considered in the same locus. Names may be slightly different in MUTAVI articles and the project model. [K3] [K4]
| Allele/combination | Typical phenotype |
|---|---|
bl+/bl+ |
green series |
blue/blue |
blue series |
yellowface_type1 + blue |
Yellowface Type I Blue |
yellowface_type2 + blue |
Yellowface Type II Blue |
goldenface + blue |
Goldenface Blue |
aqua + blue |
Aqua Blue |
turquoise + blue |
Turquoise Blue |
aqua + turquoise |
Turquoise Aqua |
Examples of dick matches:
| matching | Conclusion |
|---|---|
| Blue x Blue | %100 Blue |
| Blue x Green split Blue | %50 Blue, %50 Green split Blue |
| Green split Blue x Green split Blue | %25 Blue, %50 Green split Blue, %25 Green |
| YF2 Blue x Blue | %50 YF2 Blue, %50 Blue |
| Goldenface Blue x Blue | %50 Goldenface Blue, %50 Blue |
| Aqua Blue x Turquoise Blue | %25 Aqua Blue, %50 Turquoise/Aqua, %25 Turquoise Blue |
Notes:
Yellowface Type IThe double factor can appear as a white face in the blue series. This is one of the areas where names differ between standards. [K4]Goldenface,Aqua,Turquoise,Blue Factor I/IIAviculture standardization for may vary by source; Therefore, knowing that they are at the same locus is safer than memorizing the names. [K3] [K4]
2. Dilution locus
greywing, clearwing And dilute are alleles of the same locus. The most important practical result is greywing + clearwing combination of Full-Body Greywing is to give. [K5]
| Match / combination | Conclusion |
|---|---|
greywing/greywing |
greywing |
clearwing/clearwing |
Clearwing |
dilute/dilute |
Dilute |
greywing/clearwing |
Full-Body Greywing |
greywing/dilute |
Greywing, dilute carrier |
clearwing/dilute |
Clearwing, dilute carrier |
Examples of dick matches:
| matching | Conclusion |
|---|---|
| Greywing x Clearwing | %100 Full-Body Greywing |
| Greywing x Dilute | %100 Greywing / dilute |
| Clearwing x Dilute | %100 Clearwing / dilute |
| Greywing split Dilute x Greywing split Dilute | Greywing gives weighty results; %25 dilute may come out |
3. Ino locus
ino, texas_clearbody, pearly and in aviculture comments pallid It is considered in relation to the same gene family. MUTAVI, sex-linked clearbody ino It is discussed as an allele of the locus. [P7]
Practical combinations:
| Combination | Typical name |
|---|---|
| Ino + Green series | lutino |
| Ino + Blue series | Albino |
| Cinnamon + Ino | lacewing |
| Pallid + Ino | PallidIno / lacewing-like open phenotype |
| Texas Clearbody + Ino | Texas Clearbody visible, may carry ino |
| Texas Clearbody + Pallid | Pallid Texas Clearbody |
Most Common Combined Phenotypes
| genetic combination | The resulting phenotype |
|---|---|
blue + ino |
Albino |
yesil seri + ino |
lutino |
cinnamon + ino |
lacewing |
greywing + clearwing |
Full-Body Greywing |
recessive_pied + dutch_pied/continental clearflight tipi |
Dark-Eyed Clear |
yellowface/goldenface + blue + ino |
Creamino-like phenotypes |
violet + blue + 1 dark factor |
visual violet |
Dark-Eyed Clear MUTAVI is clearly on the subject recessive pied + Dutch/continental clearflight emphasizes unity; recessive pied + Australian dominant pied does not give the same result. [K8]
Mutation Catalog
The tables below cover all mutations supported in the project. Eslesme notu The column tells you the general rule by which you will calculate.
A. Basic series, face and tone mutations
| Mutation | Heredity | Typical color/appearance | Match note | main source |
|---|---|---|---|---|
| Blue | AR | Converts the green series to the blue series; yellow pigment disappears | AR table, Blue locus | [K1] [K3] |
| Yellowface Type I | aid | Yellow mask on blue birds; white face paradox in some double factor birds | blue locus | [K3] [K4] |
| Yellowface Type II | aid | Yellow mask and sometimes yellow spread to the body | blue locus | [K3] [K4] |
| Goldenface | aid | Stronger golden yellow on the face; intense yellow spread in blue streak | blue locus | [K3] [K4] |
| Aqua | aid | Blue-sea tone shifts towards aqua/sea green | blue locus | [K3] |
| turquoise | aid | Blue-sea tone appears richer turquoise | blue locus | [K3] |
| Blue Factor I | aid | Light greenish / intermediate blue series expression | blue locus | [K3] |
| Blue Factor II | aid | Stronger intermediate blue expression than Blue Factor I | blue locus | [K3] |
| DarkFactor | aid | in green Light -> Dark -> Olive, in blue Skyblue -> Cobalt -> Mauve |
AID table | [K1] |
| Violet | aid | Purplish tone, especially in the blue series; Very visible with 1 dark factor | AID table | [K1] [K3] |
| Gray | NAME | Grey-green in green, gray in blue | AD table | [K1] [K3] |
| anthracite | aid | Very dark cosmic tone, especially in the blue series | AID table | [K3] [K14] |
B. Dilution and melanin modifiers
| Mutation | Heredity | Typical color/appearance | Match note | main source |
|---|---|---|---|---|
| Dilute | AR | Melanin is severely reduced; The bird looks lighter overall | AR table, dilution locus | [K5] |
| greywing | AR | Wing stripes grey, body color semi-dilute | AR table, dilution locus | [K1] [K5] |
| Clearwing | AR | Wings open, body color remains bright | AR table, dilution locus | [K1] [K5] |
| cinnamon | SLR | Black melanin turns brown; warmer tone | SLR table | [K1] [K3] |
| slate | SLR | Dull blue-gray/slate tone in blue series, cheek stain becomes matte dark | SLR table | [K3] [K11] |
| blackface | Controversial | Melanin expands on the face and body, the mask darkens | Sources differ; controversial | [K3] [K9] |
| faded | AR | Washed, pale appearance; wing prints are weak | AR table | [K3] |
| Fallow (English) | AR | Red/plum eyes, warm light body tone | AR table | [K1] [K3] |
| Fallow (German) | AR | Similar to English fallow but described as a separate locus | AR table | [K3] |
| Fallow (Scottish) | AR | Bronze markings and red eyes | AR table | [K3] |
| pallid | SLR | Melanin decreases but is not completely eliminated; light bird with soft stripes | SLR table, ino family | [K3] |
| Ino | SLR | Lutino in the green series, Albino in the blue series | SLR table, ino family | [K1] [K3] [K7] |
C. Pattern, ala and clearbody mutations
| Mutation | Heredity | Typical color/appearance | Match note | main source |
|---|---|---|---|---|
| opaline | SLR | Bass and wing barring decreases, V-like pattern appears on the back | SLR table | [K1] [K3] |
| spangle | aid | Reverse wing pattern in one factor; almost solid yellow/white at double factor | AID table | [K1] [K3] |
| Recessive Pied | AR | Random light spots; eye ring behavior may differ | AR table | [K1] [K8] |
| Dominant Pied (Australian) | NAME | Clean open bands / areas on body | AD table | [K1] [K3] |
| Clearflight Pied | AR or controversial standard | Gap on flight feathers and tail, small head spot | May vary depending on source | [K1] [K8] |
| Dutch Pied | AD or AID is disputed | Irregular pattern; Clearflight-like selection on some lines | There is a difference in source | [K1] [K3] [K8] |
| Saddleback | AR | Clear saddle / V-like area on back, bass barring distinct | AR table | [K1] [K3] |
| Mottled | Controversial | Open areas that increase as the coat changes, progressive graying appearance | Sources differ | [K3] |
| Pearly | SLR | Pearl/paired edging on wing markings | SLR table, ino family | [K3] [K7] |
| Texas Clearbody | SLR | Body melanin decreases, wing lines remain dark | SLR table, ino family | [K1] [K7] |
| Dominant Clearbody | NAME | Body cleaner and lighter, wings may remain darker | AD table | [K3] [K7] |
D. Coat structure and special health-related conditions
| Mutation | Heredity | typical appearance | Match note | main source |
|---|---|---|---|---|
| Crested (Tufted) | NAME | A single or clustered hill on top | AD table; crest x crest risky | [K3] [K10] |
| Crested (Half-Circular) | NAME | semicircular hill | AD table; crest x crest risky | [K3] [K10] |
| Crested (Full-Circular) | NAME | Full circle corona peak | AD table; crest x crest risky | [K3] [K10] |
| Feather Duster | AR, pathological | Extremely long, constantly growing feathers; flight and nutrition problem | Situation to avoid mating | [K15] |
Frequently Asked Matches
Green split Blue with Blue
Blue x Green split Blue- Conclusion:
%50 Blue,%50 Green split Blue
Normal female with Cinnamon
erkek Cinnamon visual x disi normal- Male offspring:
%100 split cinnamon - Female offspring:
%100 cinnamon
Normal female with Ino
erkek Ino visual x disi normal- Male offspring:
%100 split ino - Female offspring:
%100 ino
Greywing and Clearwing
- If both parents are pure at the respective locus
Greywing x Clearwing = %100 Full-Body Greywing[K5]
Dutch Pied with Recessive Pied
Recessive Pied x Dutch Pied / clearflight tipi- In some puppies
Dark-Eyed ClearCombinations leading to the line may occur. [K8]
Ino with Cinnamon
- Pure logic of inheritance
Lacewinggoes to the phenotype. - However
cinnamonAndinoRare crossing-over results are unexpected because they are close to each other on the Z chromosomeino,cinnamonor intermediate forms may also be seen. [K6] [K12]
Risky and Controversial Areas
1. Crest x crest
In the studies compiled by MUTAVI, it is stated that crest factor is subvital and may be associated with embryo losses and neurological problems. In practice crest x crest matching should be handled with caution. [K10]
2. Blackface classification
This is one of the clearest areas of discussion:
- MUTAVI 2007 article
blackfaceforotosomal resesifsays. [K9] - Current mutation model in the project
dominantuses the comment.
In this guide blackface, marked as controversial; When making a breeding plan it is necessary to record the actual results in your own breeding line.
3. Dutch Pied and Clearflight relationship
MUTAVI argues that there may be a selective relationship between Dutch pied and continental clearflight. In contrast, in modern databases these are sometimes modeled as separate mutations. [K8]
4.Mottled
MUTAVI in the revised gene list mottle for polygenic The expression is seen; The data model in the application can treat this as a single mutation. [K3]
5. Yellowface nomenclature
Yellowface Type I, Type II, Goldenface, Aqua, Turquoise, Blue Factor I/II Their names are not exactly the same among the sources and country traditions. Here, the logic of the locus is more important than the fixed name. [K3] [K4]
Visual Reference Bank
- WBO Color Standards: the main reference for basic colors and official show varieties. [K1]
- WBO Pictorial Standard PDF: one-page visual standard summary. [K2]
- MUTAVI articles: main references for inheritance logic, allele series and historical test matches. [K4] [K5] [K6] [K7] [K8] [K9] [K10] [K11] [K12]
Conclusion
The most critical point in budgie genetics is to ask the following four questions rather than memorizing each mutation one by one:
1. This mutation AR, AD, AID or SLR ?
2. Is this mutation independent or blue, dilution or ino Allele of the same locus as?
3. Is it carried by a male or a female?
4. Does this mutation give a new named phenotype when combined with another mutation?
When you answer these four questions, you will calculate the vast majority of offspring probabilities correctly.
Resources
- [K1] WBO Color Standards, official color and variety standard: [https://www.world-budgerigar.org/colourstds.htm](https://www.world-budgerigar.org/colourstds.htm)
- [K2] WBO Color Budgerigar Pictorial Standard PDF: [https://www.world-budgerigar.org/photos15/WBO%20Standard%20for%20Colour%20Budgerigar%20Pictorial%20Ideal.pdf](https://www.world-budgerigar.org/photos15/WBO%20Standard%20for%20Colour%20Budgerigar%20Pictorial%20Ideal.pdf)
- [K3] MUTAVI, Revised List of Mutant Genes of the Budgerigar: [https://www.mutavi.info/index.php?art=symbols](https://www.mutavi.info/index.php?art=symbols)
- [K4] MUTAVI, Gene function in Yellowface Budgerigars: [https://www.mutavi.info/index.php?art=yellowface](https://www.mutavi.info/index.php?art=yellowface)
- [K5] MUTAVI, Phenotypic Effects Caused by the Multiple Allele Series of the dil-locus: [https://www.mutavi.info/index.php?art=dilute](https://www.mutavi.info/index.php?art=dilute)
- [K6] MUTAVI, Crossing-over in the Sex-chromosome of the Male Budgerigar: [https://www.mutavi.info/index.php?art=sexchrom](https://www.mutavi.info/index.php?art=sexchrom)
- [K7] MUTAVI, Genotypic and Phenotypic Aspects of the Sex-Linked clearbody: [https://www.mutavi.info/index.php?art=clearbod](https://www.mutavi.info/index.php?art=clearbod)
- [K8] MUTAVI, Recessive Pied in the Origin of Dark Eyed Clears: [https://www.mutavi.info/index.php?art=blackeye](https://www.mutavi.info/index.php?art=blackeye)
- [K9] MUTAVI, Blackface: a new mutation in the budgerigar: [https://www.mutavi.info/index.php?art=blackfa](https://www.mutavi.info/index.php?art=blackfa)
- [K10] MUTAVI, Crest: A Subvital Character in the Budgerigar: [https://www.mutavi.info/index.php?art=crested](https://www.mutavi.info/index.php?art=crested)
- [K11] MUTAVI, Description of the Slate Budgerigar: [https://www.mutavi.info/index.php?art=slate](https://www.mutavi.info/index.php?art=slate)
- [K12] MUTAVI, The lacewing: An Enigma in Budgerigar Breeding?: [https://www.mutavi.info/index.php?art=lacewing](https://www.mutavi.info/index.php?art=lacewing)
- [K13] WBO, Recognition of New Mutations: [https://world-budgerigar.org/recomutations.htm](https://world-budgerigar.org/recomutations.htm)
- [K14] WBO Noticeboard 2011, Anthracite recognition note: [https://www.world-budgerigar.org/noticeboard11.htm](https://www.world-budgerigar.org/noticeboard11.htm)
- [K15] OMIA, Feather, abnormal growth in budgerigar: [https://omia.org/OMIA002106/13146/](https://omia.org/OMIA002106/13146/)
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